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↑Olson, Storrs L. (1985). "The Fossil Record of Birds (Section X.G.5.a Sulidae)". In Farner, D.S.; King, D.S.; Parkes, K.C. (eds.). Avian Biology. Vol.8. New York: Academic Press. pp.79–238 [203–204]. hdl:10088/6553. Archived from the original on 2021-04-10. Retrieved 2017-11-22.
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↑"Sula, n.". Oxford English Dictionary (onlineed.). Oxford University Press.(Subscription or participating institution membership required.)
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↑ AviList Core Team (2025). "AviList: The Global Avian Checklist, v2025" . doi : 10.2173/avilist.v2025 . 2026年4月5日取得.